- Tassili N'Ajjer II#;
- Ahaggar II (Algeria);
- Aïr & Ténéré VIII (Niger);
- specimen collected at Armumuit oasis, Adrar Souttouf Mts, Chad (Hemmer et al. 1976);
- Djebel Bou-Hedma II* (Tunisia);
- Hai Bar Yotvata IV (Israel);
- Harrat al-Harrah IV;
- Tubayq IV;
- Mahazat as Sayed I (Saudi Arabia);
- Wahibah Sands (proposed: Oman);
- specimens collected from the Al Liwa oasis, Empty Quarter, United Arab Emirates
(M. Reza Khan in litt 1993);
- Jal az Zhor V (Kuwait);
- Moteh V (M. Moinian pers. comm.; cited in Groves 1990);
- Touran V* complex (M. Karami pers. comm.) (Iran);
- Repetek I*;
- Krasnovodsk I (Turkmenistan);
- Ustyurt I (Kazakhstan);
- Kyzylkum I (Uzbekistan);
- Registan Desert Wildlife Mgt. Reserve (proposed: Afghanistan).
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